Tissues and meristematic tissue
Std 9–10 introduced tissues simply; here we study each tissue and the internal structure of root, stem and leaf in detail.
Fresh bricks raise a new wall; likewise a group of young, dividing cells builds new parts of a plant – the meristematic tissue.
Tissue = a group of cells with a common origin and function. By ability to divide: meristematic and permanent.
In the figure : Classification of meristems: origin, position and function.
Rule 1 : Features of meristematic tissue — Young, living, dividing cells; polyhedral (isodiametric); no intercellular spaces; thin cellulose wall; dense cytoplasm; large nucleus; tiny vacuoles; high metabolism.
- Easy : Thin wall + dense cytoplasm + big nucleus = meristem.
- Medium : Cells just behind a root tip divide.
- Exam-type (CET) : Meristem cells lack a large vacuole.
Rule 2 : Classification: origin, position, function — Origin: primordial (promeristem) → primary → secondary. Position: apical (length), intercalary (near nodes, monocots), lateral (girth; e.g. cambium). Function: protoderm → epidermis; procambium → primary xylem/phloem; ground meristem → cortex, pith.
- Easy : Secondary meristem is always lateral; e.g. cork cambium.
- Medium : Intercalary tissue is short-lived; seen in grasses.
- Exam-type (CET) : Shoot apex is terminal; root apex lies behind the cap (subterminal).
Three criteria: origin, position, function; apical = height, lateral = girth.
‘Lateral meristem increases length’ is wrong (girth). Trap: ‘meristem cells have a large vacuole’ is wrong.
Memory clue : "A-L-I: Apical = Ascend, Lateral = Large girth, Intercalary = Internode".
Spot the difference
| Type | Role | Example |
|---|---|---|
| Apical | Length growth | Shoot/root tip |
| Intercalary | Node growth | Grass |
| Lateral | Girth growth | Cambium |
Practice questions
- 1What is a tissue?
- 2Two features of meristem.
- 3Role of apical meristem?
- 4Example of lateral meristem.
- 5What does protoderm form?
- 6Procambium forms?
- 7Where is intercalary meristem?
- 8Primary vs secondary meristem.
Show answers (try them yourself first)
- 1A group of cells with common origin and function.
- 2Thin wall; large nucleus.
- 3Increases length.
- 4Cambium; cork cambium.
- 5Epidermis.
- 6Primary xylem and phloem.
- 7Near nodes; mostly monocots.
- 8Primary at tips from the start; secondary arises later from permanent tissue, lateral.
Simple permanent tissues
Brick, sand and rope each do a different job; in plants parenchyma fills, collenchyma gives flexible support, sclerenchyma gives hard support.
Permanent tissue = cells that lost the ability to divide and took a fixed shape and function. Simple = one cell type: parenchyma, collenchyma, sclerenchyma.
In the figure : Three simple tissues: wall structure and function.
Rule 1 : Parenchyma and collenchyma — Parenchyma: living, thin cellulose wall, large vacuole; intercellular spaces; linked by plasmodesmata; stores food and water; photosynthesis; gas exchange. Collenchyma: living; extra cellulose and pectin at corners; no spaces; flexible support to young stems and petioles; absent in monocots and roots.
- Easy : Parenchyma = least specialised.
- Medium : Collenchyma = living mechanical tissue.
- Exam-type (CET) : Dedifferentiated parenchyma forms cambium and cork cambium.
Rule 2 : Sclerenchyma — Dead, thick lignified wall, no nucleus. Two types: fibres (long, tapering, in bundles; jute, flax, hemp) and sclereids (short, deep pits; seed coats, shells). The main supporting tissue; stiffens leaves.
- Easy : Sclerenchyma is dead; lignin gives hardness.
- Medium : Commercial fibres = sclerenchyma fibres.
- Exam-type (CET) : Sclereids arise by thickening of parenchyma.
P living filler; C living flexible; S dead hard.
‘Collenchyma is dead’ is wrong. Trap: ‘sclerenchyma has nucleus and vacuole’ is wrong.
Memory clue : "P-C-S: Plenty-space, Corner-thick, Stone-hard".
Spot the difference
| Tissue | Cell | Support |
|---|---|---|
| Parenchyma | Living, spaces | Little |
| Collenchyma | Living, corner-thick | Flexible |
| Sclerenchyma | Dead, lignin | Rigid |
Practice questions
- 1Two groups of permanent tissue.
- 2Two functions of parenchyma.
- 3Collenchyma supports?
- 4Sclerenchyma is hard due to?
- 5Two examples of fibres.
- 6Is collenchyma in monocots?
- 7Fibres vs sclereids.
- 8How does parenchyma form cambium?
Show answers (try them yourself first)
- 1Simple and complex.
- 2Food storage; photosynthesis.
- 3Young stems, leaf petiole.
- 4Lignin.
- 5Jute and flax.
- 6Usually absent.
- 7Fibres long, tapering; sclereids short, broad.
- 8Dedifferentiation (regains division).
Complex permanent tissue: xylem
Like a city water pipeline, xylem carries water and minerals from root to leaf; the pipe is made of dead cells.
Complex tissue = more than one cell type acting as one unit. Xylem (hadrome) = water- and mineral-conducting, mostly dead tissue: tracheids, vessels, xylem parenchyma, xylem fibres.
In the figure : Xylem components and thickening patterns of tracheids.
Rule 1 : Components of xylem — Tracheids: long, dead, oblique ends; 95% of gymnosperm wood, 5% of angiosperm. Vessels: wider than tracheids, end walls dissolved into a tube; angiosperms. Xylem parenchyma: the only living part; stores starch; lateral conduction. Xylem fibres: dead, lignified; mechanical support.
- Easy : Gymnosperms = tracheids; angiosperms = vessels.
- Medium : Only parenchyma is living in xylem.
- Exam-type (CET) : Vessel = many dead members joined.
Rule 2 : Thickening, protoxylem and endarch/exarch — Wall thickening: annular (rings), spiral, scalariform (ladder), reticulate, pitted (most advanced). Protoxylem = formed first, small; metaxylem = formed later, larger. Endarch = protoxylem towards pith (stem); exarch = towards periphery (root).
- Easy : Annular → spiral → scalariform → reticulate → pitted.
- Medium : Stem = endarch; root = exarch.
- Exam-type (CET) : Protoxylem small, metaxylem larger.
Xylem = dead pipe; only parenchyma is alive; stem endarch, root exarch.
‘Xylem conducts food’ is wrong (water + minerals). Trap: ‘angiosperms have only tracheids’ is wrong.
Memory clue : "X-ylem = water eXpress up; EnDarch = pith (Inner Door)".
Spot the difference
| Component | Living/Dead | Role |
|---|---|---|
| Tracheid | Dead | Conduction + support |
| Vessel | Dead | Conduction |
| Xylem parenchyma | Living | Storage |
| Xylem fibre | Dead | Support |
Practice questions
- 1What is complex tissue?
- 2Four components of xylem.
- 3Living component of xylem?
- 4Who conducts in gymnosperms?
- 5What is annular thickening?
- 6Example of endarch.
- 7Tracheid vs vessel.
- 8Protoxylem vs metaxylem.
Show answers (try them yourself first)
- 1A tissue of more than one cell type working together.
- 2Tracheid, vessel, parenchyma, fibre.
- 3Xylem parenchyma.
- 4Tracheids.
- 5Ring-like wall thickening.
- 6Stem.
- 7Vessel wider with dissolved ends; tracheid narrow and tapering.
- 8Earlier/later; small/large; annular-spiral/pitted.
Phloem, epidermal and ground tissue systems
Food made in leaves is distributed to fruits, roots and shoots; this food channel is the phloem.
Phloem (bast, leptome) = living tissue that conducts organic food: sieve tubes, companion cells, phloem parenchyma, phloem fibres. Tissue systems: epidermal, ground, vascular.
In the figure : Structure of phloem and the stomatal apparatus.
Rule 1 : Components of phloem — Sieve tubes: living tubes joined end to end; sieve plates with pores; no nucleus when mature. Companion cells: nucleus and dense cytoplasm; control the sieve tube; both arise from one cell. Phloem parenchyma: stores food, latex (absent in most monocots). Phloem fibres: dead; ropes. Sieve cells: pteridophytes and gymnosperms.
- Easy : Phloem is living (only fibres are dead).
- Medium : Sieve tube + companion cell are a pair; if one dies, so does the other.
- Exam-type (CET) : Phloem: proto (first) and meta (later).
Rule 2 : Epidermal and ground tissue systems — Epidermis: single outer layer, no gaps; cuticle saves water; root hairs in roots absorb; trichomes on stems. Stoma = pore with two guard cells (dicot: kidney; monocot: dumbbell) + subsidiary cells. Ground tissue = everything except epidermis and vascular tissue (cortex, pith, mesophyll).
- Easy : Guard cells change turgor to open/close the stoma.
- Medium : Stomatal apparatus = stoma + guard + subsidiary cells.
- Exam-type (CET) : Root hair = unicellular; trichome = multicellular.
Phloem = living food channel; guard cells = doors of the stoma.
‘Sieve tube has a nucleus’ is wrong. Trap: ‘phloem carries water’ is wrong (food).
Memory clue : "Phloem = Plant food; sieve plate = a strainer".
Spot the difference
| Component | Nature | Role |
|---|---|---|
| Sieve tube | Living, no nucleus | Food conduction |
| Companion cell | Living, nucleus | Regulation |
| Phloem fibre | Dead | Support |
Practice questions
- 1Other name for phloem?
- 2What does phloem carry?
- 3Dead part of phloem?
- 4What is a sieve plate?
- 5Function of cuticle?
- 6Trichome vs root hair.
- 7Shape of guard cells?
- 8What is ground tissue?
Show answers (try them yourself first)
- 1Bast / leptome.
- 2Organic food (source → sink).
- 3Phloem fibres.
- 4Pore-bearing end wall.
- 5Reduces water loss.
- 6Trichome multicellular on stem; root hair unicellular on root.
- 7Dicot: kidney; monocot: dumbbell.
- 8All tissue except epidermis and vascular tissue.
Vascular bundles and secondary growth
Pipes can run together or separately; the way xylem and phloem sit decides the type of vascular bundle.
Vascular bundle = a group of xylem and phloem. Secondary growth = girth increase by cambium in dicots and gymnosperms. Intrafascicular + interfascicular cambium = cambial ring.
In the figure : Types of vascular bundles and the sequence of secondary growth.
Rule 1 : Types of vascular bundles — Radial: xylem and phloem on separate radii (root). Conjoint collateral: on the same radius, xylem inside, phloem outside (stem); open = cambium present; closed = absent. Bicollateral: phloem on both sides (Cucurbitaceae). Concentric: leptocentric (phloem inside), hadrocentric (xylem inside). Amphicribral / amphivasal.
- Easy : Open = secondary growth possible; closed = not.
- Medium : Radial = root; collateral = stem.
- Exam-type (CET) : Concentric bundles are always closed.
Rule 2 : Secondary growth — Primary growth = length by apical meristem. In a dicot stem: intrafascicular cambium + dedifferentiated medullary ray cells → interfascicular cambium → cambial ring. The ring forms secondary xylem inward and secondary phloem outward; more xylem. Monocots lack cambium; exceptions: Dracaena, Agave (anomalous).
- Easy : Dedifferentiation = permanent cells regain division.
- Medium : More secondary xylem = wood.
- Exam-type (CET) : Gymnosperms also show secondary growth.
Open = cambium = grows thicker; closed = does not.
‘Monocot stem has cambium’ is wrong. Trap: ‘radial bundle in stem’ is wrong (root).
Memory clue : "Open = Operates cambium; Closed = Cannot grow".
Spot the difference
| Type | Example | Cambium |
|---|---|---|
| Radial | Root | — |
| Collateral open | Dicot stem | Yes |
| Collateral closed | Monocot stem | No |
Practice questions
- 1Where is radial bundle?
- 2Conjoint collateral means?
- 3What is in an open bundle?
- 4Bicollateral occurs in?
- 5What is leptocentric?
- 6How does a cambial ring form?
- 7Why is secondary xylem more?
- 8Why no secondary growth in monocots?
Show answers (try them yourself first)
- 1In roots.
- 2Xylem and phloem on the same radius.
- 3Cambium.
- 4Cucurbitaceae.
- 5Phloem inside, xylem outside.
- 6Intra- and interfascicular cambium join.
- 7Cambium cuts more cells inward.
- 8No cambium; bundles are closed.
Wood, cork cambium and periderm
Look at a cut tree trunk: dark centre, lighter outside, rings and bark on top – all evidence of years of secondary growth.
Wood = secondary xylem. Phellogen (cork cambium) forms phellem (cork) outside and phelloderm inside; together they are the periderm.
In the figure : Section of a trunk: heartwood, sapwood, cambium, bark and periderm.
Rule 1 : Wood: annual rings, heartwood, sapwood — In favourable seasons spring wood forms: wide lumen, thin wall, light colour, few fibres. In unfavourable seasons autumn wood: narrow, dark, thick walls, many fibres. Together they form one annual ring (clear in cold places like Kashmir). Heartwood: plugged by tyloses, oils and tannins, dead, durable; sapwood: living, conducts sap.
- Easy : One ring = one year (spring + autumn).
- Medium : Durable heartwood is used for furniture.
- Exam-type (CET) : Tyloses plug the xylem tubes.
Rule 2 : Cork cambium, periderm and bark — When the epidermis ruptures, outer cortex cells become meristematic as phellogen. Outside: cork (phellem), waterproof due to suberin; inside: phelloderm (secondary cortex). Bark = everything outside the vascular cambium. Lenticels = gas-exchange pores in the bark. Roots too show secondary growth.
- Easy : Periderm = phellogen + phellem + phelloderm.
- Medium : Cork is dead; suberin makes it waterproof.
- Exam-type (CET) : Bark is a non-technical term for all outer tissues.
Spring = wide light; autumn = narrow dark; periderm = P-P-P.
‘Phellem forms inside’ is wrong (outside). Trap: ‘heartwood is living’ is wrong.
Memory clue : "Phellem = Perimeter (outside); Phelloderm = Deep (inside)".
Spot the difference
| Point | Spring wood | Autumn wood |
|---|---|---|
| Lumen | Wide | Narrow |
| Colour | Light | Dark |
| Density | Low | High |
Practice questions
- 1What forms an annual ring?
- 2Other name for heartwood?
- 3Function of sapwood?
- 4What are tyloses?
- 5Job of phellogen?
- 6What is a lenticel?
- 7Why faint rings in Maharashtra?
- 8Why is cork waterproof?
Show answers (try them yourself first)
- 1Spring and autumn wood.
- 2Duramen.
- 3Conducting sap.
- 4Parenchyma outgrowths that plug xylem tubes.
- 5Forms cork and phelloderm.
- 6Gas-exchange pore on bark.
- 7Less seasonal difference.
- 8Due to suberin.
Anatomy of root, stem and leaf
Under the microscope a section is identified like a puzzle: bundles in a ring = dicot; scattered = monocot.
Anatomy = internal structure. T.S. of root, stem and leaf differs in dicots and monocots; leaf: dorsiventral (dicot) and isobilateral (monocot).
In the figure : Dicot and monocot stems, leaf layers and a root-stem-leaf comparison.
Rule 1 : Root and stem — Dicot root: epiblema (root hairs) → cortex → exodermis → endodermis (Casparian strip, passage cells) → pericycle → stele; 2–6 xylem, exarch; small pith; later secondary growth. Monocot root: more than 6 xylem (polyarch); large pith; no secondary growth. Dicot stem: epidermis (trichomes), collenchymatous hypodermis, cortex, endodermis (starch sheath), pericycle (hard bast), bundles in a ring, collateral open, endarch; pith present. Monocot stem: sclerenchymatous hypodermis; scattered closed bundles with bundle sheath; no pith; lysigenous cavity.
- Easy : Endodermis: Casparian strip (suberin) regulates water.
- Medium : Root is exarch; stem is endarch.
- Exam-type (CET) : Monocot stem: scattered closed bundles.
Rule 2 : Leaf — Dorsiventral (dicot): upper epidermis (few stomata) → palisade (long, many chloroplasts) → spongy (loose, spaces) → lower epidermis (more stomata, sub-stomatal chamber). Bundles closed; xylem above, phloem below; bundle sheath. Isobilateral (monocot): stomata on both sides; mesophyll not differentiated; parallel veins; bulliform cells help rolling.
- Easy : No cambium in leaves → no secondary growth.
- Medium : Palisade is above and makes food.
- Exam-type (CET) : Amphistomatic = stomata on both surfaces.
Dicot = ring, pith; monocot = scattered, no pith; root polyarch > 6.
‘Monocot stem has cambium’ is wrong. Trap: ‘dicot leaf stomata only above’ is wrong (more below).
Memory clue : "Mono = Many scattered; Di = Disciplined ring".
Spot the difference
| Point | Dicot | Monocot |
|---|---|---|
| Stem bundles | Ring, open | Scattered, closed |
| Root xylem | 2–6 | More than 6 |
| Leaf | Dorsiventral | Isobilateral |
Practice questions
- 1What is epiblema?
- 2Casparian strip is made of?
- 3What is polyarch?
- 4Bundles in dicot stem?
- 5Is there pith in monocot stem?
- 6Where is palisade parenchyma?
- 7Function of bulliform cells?
- 8Dicot vs monocot leaf.
Show answers (try them yourself first)
- 1Outermost root layer with root hairs.
- 2Suberin; in endodermal walls.
- 3More than six xylem groups (monocot root).
- 4Ring, conjoint, collateral, open, endarch.
- 5No.
- 6Below upper epidermis.
- 7Help leaf rolling.
- 8Dorsiventral/isobilateral; stomata mostly below/both; palisade-spongy/none.
Quick revision of the whole lesson
| Concept | Key fact |
|---|---|
| Meristem | Apical = length; lateral = girth |
| Simple tissues | P living; C living; S dead (lignin) |
| Xylem | Dead; water; tracheid, vessel; endarch/exarch |
| Phloem | Living; food; sieve tube + companion |
| Bundles | Radial; collateral open/closed; concentric |
| Secondary growth | Cambial ring; periderm = P-P-P |
| Anatomy | Dicot ring; monocot scattered |
🎯 Most important points for the exam
- Classification of meristems.
- Parenchyma, collenchyma, sclerenchyma comparison.
- Xylem vs phloem.
- Types of vascular bundles.
- Secondary growth, annual ring, periderm.
- T.S. of dicot/monocot root, stem, leaf.
Understand the concept → draw the figure → memorise the key points → solve questions yourself.