Biodiversity: levels and how many species
A single mango orchard has many varieties of mango, many kinds of insects and birds, and soil full of microbes. Variety exists at every level.
Biodiversity is the sum total of variety at all levels of biological organisation. The three important levels are genetic diversity (variation within a species), species diversity (variety of species in a region) and ecological diversity (variety of ecosystems). More than 1.5 million species have been described; Robert May estimated the global total at about 7 million, so only about 22 per cent are known.
In the figure : Conservation aims to protect all three levels.
In the figure : Only about 22 per cent of the estimated species have been described.
Rule 1: Levels with examples — Genetic diversity: the medicinal plant Rauwolfia vomitoria shows variation in the potency of its chemical (reserpine) in different Himalayan ranges; India has over 50,000 genetically different strains of rice and 1,000 of mango. Species diversity: the Western Ghats have more amphibian species than the Eastern Ghats. Ecological diversity: India, with deserts, rain forests, mangroves, coral reefs, wetlands, estuaries and alpine meadows, has a greater ecosystem diversity than a Scandinavian country such as Norway.
- Easy : Many rice varieties show genetic diversity.
- Medium : Western Ghats vs Eastern Ghats amphibians show species diversity.
- Exam-type : Q: What type of diversity is shown by 1,000 varieties of mango? A: Genetic diversity.
Rule 2: How many species? — More than 70 per cent of the described species are animals, and about 70 per cent of those are insects. Fungi have more species than all vertebrates together. India has about 45,000 plant species and about twice as many animal species, and is one of the 12 mega-diversity countries. Many tropical species are still unknown; prokaryote counts are very uncertain.
- Easy : Insects are the largest group of animals.
- Medium : India is a mega-diversity country.
- Exam-type : Q: What per cent of the estimated species is described? A: About 22 per cent.
Three levels: genetic, species, ecological. Described: > 1.5 million; estimated: about 7 million (May).
Mixing genetic and species diversity. Different varieties of one species = genetic diversity.
Memory clue: G-S-E — Genes, Species, Ecosystems.
Spot the difference
| Point | Genetic diversity | Species diversity |
|---|---|---|
| Level | Within a species | Between species |
| Example | Many rice varieties | Many amphibian species |
| Region | Any one species | A whole area |
Practice questions
- 1Define biodiversity.
- 2Name the three levels of biodiversity.
- 3Who estimated 7 million species on Earth?
- 4Which group has the most described species among animals?
- 5What is India's special status in terms of biodiversity?
- 6What is ecological diversity?
- 7If 1.5 million of an estimated 7 million species are described, what per cent is known?
- 8Why is the figure of 7 million only an estimate?
Show answers (try them yourself first)
- 1The variety of life at all levels of biological organisation.
- 2Genetic, species and ecological diversity.
- 3Robert May.
- 4Insects.
- 5It is one of the 12 mega-diversity countries.
- 6The variety of ecosystems in a region.
- 7About 21–22 per cent.
- 8Many tropical species and microbes are undiscovered and not counted.
Patterns of biodiversity
Count the bird species in a tropical forest and in a cold northern town; the forest wins easily.
Species diversity shows a latitudinal gradient: it decreases from the equator towards the poles. The tropics (23.5° N to 23.5° S) have the most species. Another pattern is the species-area relationship of Alexander von Humboldt: richness rises with area but only up to a limit, and on a log scale it is a straight line, log S = log C + Z log A, where S is species richness, A is area, Z is the slope and C is the Y-intercept.
In the figure : Bird species fall sharply with latitude.
In the figure : Larger areas show a steeper slope.
Rule 1: Why are the tropics richer? — Three main ideas: (a) tropics had more evolutionary time, since temperate areas were repeatedly glaciated; (b) tropical environments are less seasonal and more constant and predictable, which promotes niche specialisation; (c) more solar energy gives higher productivity and so supports more species.
- Easy : Tropical climate is more stable.
- Medium : Glaciation affected temperate regions.
- Exam-type : Q: Name two reasons for high tropical diversity. A: More evolutionary time and constant environment (or more solar energy).
Rule 2: Species-area relationship — Humboldt found that within a region species richness increases with area, up to a limit. On a log-log graph it is a straight line. For small regions Z is about 0.1 to 0.2 whatever the group or region. For very large areas such as whole continents, Z is steeper, 0.6 to 1.2.
- Easy : Larger area: more species.
- Medium : On a log scale the graph is a straight line.
- Exam-type : Q: What is the usual range of Z for small regions? A: 0.1 to 0.2.
Gradient: more species near the equator. Species-area: S rises with A; Z = 0.1–0.2 (regions), 0.6–1.2 (continents).
Thinking the species-area graph is a straight line on a normal scale. It is a rectangular hyperbola; it is a straight line only on log scales.
Memory clue: Z is small, area is small — Z is 0.1–0.2 for regions, much steeper for continents.
Spot the difference
| Point | Latitudinal gradient | Species-area relationship |
|---|---|---|
| Variable | Latitude | Area |
| Trend | Richness falls towards poles | Richness rises with area |
| Shown by | Tropics vs poles | Straight line on log scale |
Practice questions
- 1What is the latitudinal gradient?
- 2Which region has the most species?
- 3Who studied the species-area relationship?
- 4In log S = log C + Z log A, what does Z stand for?
- 5Give the usual Z value for small regions.
- 6Give the Z value range for whole continents.
- 7Which has more bird species, Colombia or Greenland?
- 8Why does a stable tropical climate help diversity?
Show answers (try them yourself first)
- 1Species diversity decreases from the equator towards the poles.
- 2The tropics.
- 3Alexander von Humboldt.
- 4The slope of the line.
- 50.1 to 0.2.
- 60.6 to 1.2.
- 7Colombia (about 1,400 against 56).
- 8It allows niche specialisation and more species coexist.
Importance of diversity and loss of biodiversity
A mixed forest survives a drought better than a plantation of one tree. Variety acts like insurance.
Ecologists believe communities with more species tend to be more stable, to have higher productivity and to resist invasions by alien species. In the rivet popper hypothesis of Paul Ehrlich, species are like rivets of an aircraft: losing a few is tolerable but the loss of many weakens the whole. Life on Earth has seen five mass extinctions; the present sixth extinction is 100 to 1,000 times faster and is caused by human activities.
In the figure : Amphibians and gymnosperms are the most threatened groups.
Rule 1: Why diversity matters — Biodiversity loss in a region may cause (a) a decline in plant production, (b) lower resistance to drought and other disturbances and (c) more variability in processes such as water use and pest and disease cycles.
- Easy : More species: more stability.
- Medium : Fewer species: lower resistance.
- Exam-type : Q: State the rivet popper hypothesis. A: Loss of a few species is tolerable, but the loss of many weakens an ecosystem like removing rivets from an aircraft.
Rule 2: Extinction facts — IUCN Red List (2004): 784 species became extinct in 500 years. Examples: dodo (Mauritius), quagga (Africa), thylacine (Australia), Steller's sea cow (Russia), and three tiger subspecies (Bali, Javan, Caspian). Over 15,500 species face extinction. Past mass extinctions numbered five. At present trends, nearly half of all species could be lost in 100 years.
- Easy : Dodo is an extinct bird of Mauritius.
- Medium : The thylacine lived in Australia.
- Exam-type : Q: How many mass extinctions occurred earlier? A: Five.
Sixth extinction = human-caused, 100–1,000 times faster than before. IUCN: 784 species extinct in 500 years.
Saying the earlier mass extinctions were also caused by humans. They were natural; only the present one is human-driven.
Memory clue: 5 then 6 — Five natural mass extinctions, the sixth is now.
Spot the difference
| Point | Earlier mass extinctions | Sixth extinction |
|---|---|---|
| Cause | Natural events | Human activities |
| Rate | Slow | 100–1,000 times faster |
| Number | Five | Ongoing |
Practice questions
- 1Name an extinct bird of Mauritius.
- 2Which animal became extinct in Australia?
- 3Who proposed the rivet popper hypothesis?
- 4How many mass extinctions happened before humans?
- 5Which group has the highest per cent threatened among the four listed?
- 6What does IUCN stand for?
- 7How fast is the present extinction rate compared to the past?
- 8Give two effects of biodiversity loss on an ecosystem.
Show answers (try them yourself first)
- 1The dodo.
- 2The thylacine.
- 3Paul Ehrlich.
- 4Five.
- 5Amphibians (32 per cent).
- 6International Union for Conservation of Nature and Natural Resources.
- 7100 to 1,000 times faster.
- 8Lower plant production and lower resistance to droughts or disturbances.
Causes of biodiversity loss: the Evil Quartet
A forest cut for a highway, a fish dropped in a lake, a hunter with too many licences: each takes away species in a different way.
The four major causes of biodiversity loss, called the Evil Quartet, are habitat loss and fragmentation, over-exploitation, alien species invasions and co-extinctions.
In the figure : Habitat loss is the most important cause.
Rule 1: The four causes — Habitat loss: rain forests once covered 14 per cent of the land and now cover about 6 per cent; the Amazon is cleared for soya and cattle. Fragmentation harms animals needing large territories and migratory animals. Over-exploitation: Steller's sea cow and passenger pigeon; over-fishing. Alien species: Nile perch in Lake Victoria wiped out over 200 cichlid species; Parthenium, Lantana and water hyacinth are weeds in India; African catfish Clarias gariepinus threatens native catfish. Co-extinction: parasites of an extinct host and the plant–pollinator pair.
- Easy : Passenger pigeon: over-exploitation.
- Medium : Nile perch: alien species.
- Exam-type : Q: Which is the most important cause? A: Habitat loss and fragmentation.
Rule 2: Linking to causes — Identify the cause by the story. Forest broken into patches → fragmentation. Fish stock collapses from heavy catch → over-exploitation. An introduced weed spreading → invasion. An orchid vanishes after its only pollinator disappears → co-extinction.
- Easy : Weed spreading over fields is invasion.
- Medium : Orchid and its pollinator is co-extinction.
- Exam-type : Q: Fish parasites vanish after the host dies. Which cause? A: Co-extinction.
Evil Quartet: habitat loss, over-exploitation, alien invasions, co-extinctions. Habitat loss is No. 1.
Thinking alien species are always harmful. Only those that become invasive cause losses.
Memory clue: H-O-A-C — Habitat, Overuse, Aliens, Co-extinction.
Spot the difference
| Point | Over-exploitation | Alien invasion |
|---|---|---|
| Cause | Excess harvesting | Introduced species spreads |
| Example | Passenger pigeon | Nile perch in Lake Victoria |
| Result | Population crash | Native species decline |
Practice questions
- 1Name the four causes of biodiversity loss.
- 2Which cause is the most important?
- 3Give an example of an invasive weed in India.
- 4Which fish caused extinction of cichlids in Lake Victoria?
- 5What is co-extinction?
- 6Give one over-exploited species.
- 7Rain forests covered 14 per cent of land earlier and 6 per cent now. How much of the land cover did they lose?
- 8Why are migratory animals affected by fragmentation?
Show answers (try them yourself first)
- 1Habitat loss and fragmentation, over-exploitation, alien species invasions and co-extinctions.
- 2Habitat loss and fragmentation.
- 3Parthenium, Lantana or water hyacinth.
- 4Nile perch.
- 5Extinction of species dependent on an extinct species.
- 6Steller's sea cow or passenger pigeon.
- 78 percentage points of land surface.
- 8They cannot move through broken habitat, so populations decline.
Why conserve biodiversity?
Neem, tulsi, turmeric, bees and mangroves all help us daily. Some give direct products and some give silent services.
Reasons for conservation fall into three groups. Narrowly utilitarian: direct benefits such as food, fibre, firewood, medicines and industrial products. Broadly utilitarian: ecosystem services such as oxygen production and pollination. Ethical: every species has an intrinsic value and we have a moral duty to pass the legacy to future generations.
In the figure : Each reason adds a different kind of value.
Rule 1: Utilitarian arguments — More than 25 per cent of drugs sold worldwide come from plants, and 25,000 plant species are used in traditional medicines. Bioprospecting explores biodiversity for economically useful products. The Amazon forest produces about 20 per cent of the oxygen in the atmosphere. Pollinators (bees, birds, bats) enable fruit and seed formation. Aesthetic pleasure is an intangible benefit.
- Easy : Plant-based drugs are a direct benefit.
- Medium : Pollination is an ecosystem service.
- Exam-type : Q: What is bioprospecting? A: Exploring molecular, genetic and species diversity for products of economic value.
Rule 2: Ethical argument — Species have a right to exist regardless of their use to humans. We share the planet with millions of plants, animals and microbes and owe them care.
- Easy : A species without economic use still matters.
- Medium : It is a moral duty.
- Exam-type : Q: What is the ethical argument? A: Every species has an intrinsic value and we must protect it.
Three reasons: narrowly utilitarian, broadly utilitarian, ethical. Amazon gives about 20 per cent of the world's oxygen.
Saying only economic value matters. The ethical argument is about intrinsic value.
Memory clue: N-B-E — Narrow (products), Broad (services), Ethical (right to live).
Spot the difference
| Point | Narrowly utilitarian | Broadly utilitarian |
|---|---|---|
| Benefit | Direct products | Ecosystem services |
| Example | Medicines, fibre | Pollination, oxygen |
| Value | Easy to price | Hard to price |
Practice questions
- 1Name the three reasons for conserving biodiversity.
- 2What per cent of the Earth's oxygen comes from the Amazon forest?
- 3What is bioprospecting?
- 4Give one ecosystem service of bees.
- 5What per cent of drugs come from plants?
- 6Which argument supports intrinsic value?
- 7A tribal community uses 25 plants for medicine. Which argument does this support?
- 8Why is the broadly utilitarian argument hard to put a price on?
Show answers (try them yourself first)
- 1Narrowly utilitarian, broadly utilitarian and ethical.
- 2About 20 per cent.
- 3Exploring biodiversity for products of economic value.
- 4Pollination.
- 5More than 25 per cent.
- 6The ethical argument.
- 7Narrowly utilitarian.
- 8Services like clean air and pollination are benefits not sold in markets.
How to conserve biodiversity
To save the tiger, we protect the whole forest. To save an animal on the brink, we may need a safe enclosure or a seed bank.
In situ conservation protects species on site, in their natural habitat. Ex situ conservation protects them off site, in zoos, botanical gardens, gene banks and similar places. Biodiversity hotspots are regions with very high species richness and high endemism, and they are also regions of accelerated habitat loss.
In the figure : In situ keeps the whole ecosystem safe; ex situ is for species at very high risk.
Rule 1: Hotspots and in situ — There are 34 hotspots in the world (initially 25, then 9 more). Three cover India: Western Ghats and Sri Lanka, Indo-Burma and Himalaya. They cover less than 2 per cent of the land, yet strict protection could cut the ongoing mass extinctions by almost 30 per cent. India has 14 biosphere reserves, 90 national parks and 448 sanctuaries, plus sacred groves in Meghalaya (Khasi and Jaintia Hills), Aravalli Hills, the Western Ghats and Madhya Pradesh.
- Easy : Hotspots: less than 2 per cent of land.
- Medium : Sacred groves are in Meghalaya.
- Exam-type : Q: Name the three hotspots of India. A: Western Ghats and Sri Lanka, Indo-Burma, Himalaya.
Rule 2: Ex situ and international efforts — Ex situ methods: zoological parks, botanical gardens, wildlife safari parks, cryopreservation of gametes, in vitro fertilisation, tissue culture and seed banks. The Convention on Biological Diversity (Earth Summit) was held at Rio de Janeiro in 1992. The 2002 World Summit on Sustainable Development in Johannesburg: 190 countries pledged a major reduction in biodiversity loss by 2010.
- Easy : Seed bank is an ex situ method.
- Medium : Earth Summit: Rio, 1992.
- Exam-type : Q: Where and when was the Earth Summit? A: Rio de Janeiro, 1992.
34 hotspots in the world; 3 in India. In situ = on site; ex situ = off site.
Calling a zoo in situ conservation. A zoo is off site, so it is ex situ.
Memory clue: In → inside habitat — In situ = inside the natural home.
Spot the difference
| Point | In situ | Ex situ |
|---|---|---|
| Place | Natural habitat | Outside the habitat |
| Example | National park | Zoo, seed bank |
| Best for | Protecting a whole ecosystem | Species at very high risk |
Practice questions
- 1Define in situ conservation.
- 2Give two ex situ methods.
- 3How many hotspots are there in the world?
- 4Name one hotspot in India.
- 5What is endemism?
- 6Where are sacred groves found?
- 7Initially 25 hotspots were identified and 9 added later. What is the total?
- 8Why are hotspots given priority in conservation?
Show answers (try them yourself first)
- 1Protecting species in their natural habitat.
- 2Zoos and seed banks (or cryopreservation).
- 334.
- 4Western Ghats and Sri Lanka, Indo-Burma or Himalaya.
- 5Species being confined to one region and not found elsewhere.
- 6Meghalaya and the Aravalli and Western Ghats regions, among others.
- 734.
- 8They hold very high species richness and endemism but are losing habitat fast.